^ indicates undergraduate student, * graduate student, and ** postdoctoral fellow.


89. Vollset K, & M Krkosek, 2001. Delayed maturity does not offset negative impact afflicted by ectoparasitism in Salmon. Theoretical Ecology, in press.

88. Krkosek M, *Jarvis-Cross M, *Wadhawan K, **Berry I, **Soucy JPR, *Bodner K, *Greiner A, , *Krichel L, *Penk S, *Vargas-Soto J, *Shea D, **Tekwa E, Mideo N, Molnar P,  2021. Establishment, contagiousness, and initial spread of SARS-CoV-2 in Canada. FACETS, 6, 180-194.


87. *Kirk D, **Luijckx P, **Jones N, *Krichel L, ^Pencer C, Molnar P, & Krkosek M, 2020. Experimental evidence of warming-induced disease emergence and its prediction by a trait-based mechanistic model. Proceedings of the Royal Society B, 287, 20201526.

86. *Godwin S, Krkosek, M, Reynolds J, **Bateman A., 2020. Sea-louse abundance on salmon farms in relation to parasite-control policy and climate change. ICES Journal of Marine Science, fsaa173.

85. *Shea D, **Bateman A, Li S, Tabata A, Schulze A, **Mordecai G, *Ogston L, Volpe J, Frazer L, Connors B, Miller K, Short S, & Krkosek M., 2020. Environmental DNA (eDNA) from multiple pathogens is elevated near active Atlantic salmon farms. Proceedings of the Royal Society B. 287: 20202010

84. *Godwin S, Krkosek M, Reynolds J, & Bateman A., 2020. Bias in self-reported parasite data from the salmon farming industry. Ecological Applications. 31, e02226.
News coverage in CBC and the Narwhal

83. ^Brookson C, Krkosek M, Hunt B, Johnson B, **Rogers L, & **Godwin S, 2020. Differential infestation of juvenile Pacific salmon by parasitic sea lice in British Columbia, Canada. Canadian Journal of Fisheries and Aquatic Sciences, 77, 1960-1968.
News coverage in Hakai Magazine

82. **Peacock S, Krkosek M, Lewis M & Molnar P, 2020. A unifying framework for the transient parasite dynamics of migratory hosts. Proceedings of the National Academy of Sciences of the USA 117, 10897-10903.

81. ^Zukowski N, *Kirk D, *Wadhawan K, *Shea D & Krkosek M, 2020. Predators can influence the host-parasite dynamics of their prey via non-consumptive effects. Ecology and Evolution 10, 6714–6722.

80. Behringer D, Wood C, Krkosek M & Bushek D, 2020. Disease in fisheries and aquaculture. In Behringer D, K Lafferty, and B Silliman, Marine Disease Ecology. Oxford University Press, Oxford, UK.

79. **Bateman A, **Peacock S, Krkosek M, & Lewis M, 2020. Migratory hosts  can maintain the high-dose refuge effect in a structured host-parasite system: the case of sea lice and salmon. Evolutionary Applications, 13, 2521-2535.


78. **Peacock S, Krkosek M, Bateman A, & Lewis M, 2019. Estimation of spatiotemporal transmission dynamics and analysis of management scenarios for sea lice of farmed and wild salmon. Canadian Journal of Fisheries and Aquatic Sciences 77, 55-68.

77. **Peacock S, ** Bateman A, Connors B, * Godwin S, Lewis M, & Krkosek M, 2019. Ecology of a marine ectoparasite in farmed and wild salmon. In Wilson, Fenton, Tompkins (eds) Wildlife Disease Ecology. Cambridge University Press, Cambridge, UK.

76. **Darling E, …, Fortin MJ, Krkosek M, & Mouillot D, 2019. Social–environmental drivers inform strategic management of coral reefs in the Anthropocene. Nature Ecology and Evolution 3, 1341–1350.

75. *Kirk D, **Luijckx P, ^Stanic A, Krkosek M, 2019. Predicting the thermal and allometric dependencies of disease transmission via the metabolic theory of ecology. American Naturalist 193, 661-676.


74. ^Atkinson E, **Bateman A, Dill L, Krkosek M, Reynolds J, & *Godwin S. Oust the louse: Leaping behaviour removes sea lice from wild juvenile sockeye salmon. Journal of Fish Biology 93, 163-271.
News coverage in Science

73. **Searcy C, Gilbert B, Krkosek M, Rowe L, & McCauley S, 2018. Positive correlation between dispersal and body size in green frogs (Rana clamitans) naturally colonizing and experimental landscape. Canadian Journal of Zoology 96, 1378-1384

72. **Bateman A, Ozgul A, Krkosek M, & Clutton-Brock T., 2018. Matrix models of hierarchical demography: Linking group- and population-level dynamics in cooperative breeders. American Naturalist 192, 188-203.

71. ^Phillips J, **Peacock S, *Bartlett M, **Bateman A, Lewis M, & Krkosek M., 2018. An asymmetric producer-scrounger game: body size and the social foraging behavior of coho salmon. Theoretical Ecology 11, 417-431.

70. *Rogers L, Salomon A, Connors B, & Krkosek M., 2018. Collapse, tipping points, and spatial demographic structure arising from the adopted migrant life history. American Naturalist 192, 49-61.
News coverage in The Varcity

69. *Kirk D, *Jones N, **Peacock S, ^Phillips J, Molnar P, Krkosek M, **Luijckx P, 2018. Empirical evidence that metabolic theory describes the temperature dependency of within-host parasite dynamics. PLoS Biology 16, e2004608.
Highlighted by a Primer in PLoS Biology

68. *Dallas, T. M. Krkosek, & J. Drake, 2018. Pathogen invasion thresholds in a Daphnia-microparasite system. Royal Society Open Science 5, 171975.

67. *Godwin S, Krkosek M, Reynolds R, *Rogers, L & Dill L., 2018. Heavy sea louse infection is associated with decreased foraging success in wild juvenile sockeye salmon. Canadian Journal of Fisheries and Aquatic Sciences 75, 1587-1595.


66. *Meyer, S., L. Chilvers, B. Robertson, & M. Krkosek, 2017 Marine mammal population decline linked to obscured bycatch. Proceedings of the National Academy of Sciences of the USA, 114: 11781–11786.
News coverage in Hakai Magazine, Science Daily, NZ Herald.

65. **Vollset, K., I. Dohoo, O. Karlsen, E. Halttunen, B. Kvamme, B. Finstad, V. Wennevik, O. Diserud, A. **Bateman, K. Friedland, S. Mahlum, C. Jergensen, L. Qviller, M. Krkosek, A. Atland, B. Barlaup, 2017. Food for thought: Disentangling the role of sea lice on the marine survival of Atlantic salmon. ICES Journal of Marine Science. 75, 50–60.

64. *Kreitzman, M., J. *Ashander, J. *Driscoll, A. **Bateman, K. Chan, M. Lewis, & M. Krkosek, 2017. An evolutionary ecosystem service: wild salmon sustain the effectiveness of parasite control on salmon farms. Conservation Letters, 11, 1-13.

63. Krkosek, M., 2017. Population biology of infectious diseases shared by wild and farmed fish. Canadian Journal of Fisheries and Aquatic Sciences, 74, 620-628.
Editors Choice in CJFAS and highlighted in the Dispatch section of Frontiers in Ecology and the Environment

62. *Godwin, S., L. Dill, M. Krkosek, M. *Price, and J. Reynolds, 2017. Reduced growth of wild juvenile sockeye salmon associated with sea louse infection. Journal of Fish Biology, 91 41-57.
News coverage in CBC


61. *Peacock, S., M. Krkosek, M. Lewis, & S. Lele, 2016. Study design and parameter estimability for spatial and temporal ecological models. Ecology and Evolution, 7, 762–770.

60. **Bateman, A., S. *Peacock, B. Connors, ^D. Berg, Z. Polk, M. Krkosek, & A. Morton, 2016. Recent failure in control of sea lice epizootics on salmon in the Broughton Archipelago, British Columbia. Canadian Journal of Fisheries and Aquatic Sciences, 73, 1164-1172.
News coverage by the Vancouver Sun and the Tyee.

59. *Meyer, S., B. Robertson, L. Chilvers, & M. Krkosek, 2016. Reply to Comment on “Population dynamics reveal conservation priorities of the threatened New Zealand sea lion Phocarctos hookeri” by Middleton and Breen (2016). Marine Biology, 163, 113.

58. *Peacock, S., A. **Bateman, M. Krkosek, B. Connors, S. *Rogers, L. Portner, Z. Polk, C. Webb, & A. Morton, 2016. Sea-louse parasites on juvenile wild salmon in the Broughton Archipelago, British Columbia, Canada. Ecology, 97: 1887 (data paper).

57. *Peacock, S., A. **Bateman, M. Krkosek, & M. Lewis, 2016. The dynamics of coupled populations subject to control. Theoretical Ecology, 9, 365.

56. **Groner, M.L., L. *Rogers, A. **Bateman, B. Connors, L.N. Frazer, S. *Godwin, M. Krkosek, M. Lewis, S. *Peacock, E. Rees, C. Revie, & U. *Schlagel 2016. Lessons from sea louse and salmon epidemiology. Philosophical Transactions of the Royal Society B, 371, 20150203.


55. Closs. G., M. Krkosek, & J. Olden, 2015. Conservation of Freshwater Fishes. Cambridge University Press. Cambridge, UK.

54. Closs G, Krkosek M, & Olden J, 2014. Synthesis – what is the future of freshwater fishes? In Conservation of Freshwater Fishes. Editors: G. Closs, M. Krkosek, & J. Olden. Cambridge University Press, Cambridge.

53. Krkosek, M., & R. Poulin, 2015. Infectious disease and the conservation of freshwater fish. In Conservation of Freshwater Fishes. Editors: G. Closs, M. Krkosek, & J. Olden. Cambridge University Press, Cambridge.

52. **Vollset, K., R. Krontveit, P. Jansen, B. Finstad, B. Barlaup, O. Skilbrei, M. Krkosek, P. Romunstad, A. Aunsmo, A. Jansen, & I. Doohoo, 2015. Impacts of parasites on marine survival of Atlantic salmon: a meta-analysis. Fish and Fisheries 7, 91-113.

51. *Peacock, S., M. Krkosek, A. **Bateman & M. Lewis, 2015. Parasite-mediated release from predation in a juvenile salmon food web. Ecosphere. 6:art264.

50. *Meyer, S. B. Roberson, L. Chilvers, and M. Krkosek, 2015. Population dynamics reveal life history based priorities for the conservation management of the imperilled New Zealand sea lion. Marine Biology. 162, 1587-1596.
News coverage by Radio New Zealand and Otago Daily Times.

49. *Meyer, S., A. *Davidson, M. Krkosek, & B. Robertson, 2015. Comment on”Current bycatch levels in Auckland Islands trawl fisheries unlikely to be driving New Zealand sea lion (Phocarctos hookeri) population decline”. Aquatic Conservation: Marine and Freshwater Ecosystems 4, 584-586.

48. *Godwin, S., L. Dill, J. Reynolds, & M. Krkosek 2015. Sea lice, sockeye salmon, and foraging competition: lousy fish are lousy competitors. Canadian Journal of Fisheries and Aquatic Sciences. 72, 1113-1120.
News coverage by the Vancouver Sun and the Globe and Mail.

47. **Rees, E., S. St-Hilaire, S. Jones, M. Krkosek, S. DeDominicis, M. Foreman, T. *Patanasatienkul & C. Revie, 2015. Spatial patterns of sea lice infection among wild and captive salmon in western Canada. Landscape Ecology. 30, 989-1004.

46. **Bateman, A., M. Neubert, M. Krkosek, & M. Lewis, 2015. Generational spreading speed and the dynamics of ecological invasions. American Naturalist 186, 362-375.

45. Randhawa, H., R. Poulin, & M. Krkosek 2015. Increasing rate of species discovery in sharks coincides with sharp population declines: implications for biodiversity. Ecography, 38, 96-107.
Cover story

44. Tompkins, D.M., S. Carver, M.E. Jones, M. Krkosek, L. Skerratt, 2015. Emerging infectious diseases of wildlife: a critical perspective. Trends in Parasitology 31, 149-159.


43. Krkosek, M. & J. Drake, 2014. On signals of phase transitions in salmon population dynamics. Proceedings of the Royal Society B. 281, 20133221.

42. *Sommerville, G., M. Krkosek, & C. Hepburn, 2014. A matrix model and elasticity analysis for New Zealand’s blackfoot paua Haliotis iris. Fisheries Research, 151, 158-168.

41. **Bird, T.J., A.E. Bates, J.S. Lefcheck, N.Hil, S. Wotherspoon, M. Krkosek, R.D. Stuart-Smith, J.F. Stuart-Smith, G. Pecl, G. Edgar, R.J. Thomson, N. Barrett, & S. Frusher, 2014. Statistical solutions for error and bias in global citizen science datasets. Biological Conservation, 173, 144-154.

40. *Peacock, S., B. Connors, M. Krkosek, J. Irvine, & M. Lewis. 2014. Can reduced predation offset negative effects of sea louse parasites on chum salmon? Proceedings of the Royal Society B. 281, 20132913.

39. *Akbaripasand, A., M. Krkosek, P. M. Lokman, & G. Closs, 2014. Does social status within a dominance hierarchy mediate individual growth, residency and relocation? Oecelogia, 176, 771-779.

38. Krkosek, M., C. Revie, B. Finstad, & C. Todd, 2014 Comment on Jackson et al. “Impact of Lepeophtheirus salmonis infestations on migrating Atlantic salmon, Salmo salar L., smolts at eight locations in Ireland with an analysis of lice-induced marine mortality” Journal of Fish Diseases. In press.
News coverage by the Scotland Herald and the Irish Times.


37. **Studer, A., M. *Widmann, R. Poulin, & M. Krkosek, 2013. Large scale patterns of trematode parasitism in a bivalve host: no evidence for a latitudinal gradient in infection levels. Marine Ecology Progress Series. 491, 125-135.

36. Krkosek, M., J. *Ashander, L.N. Frazer, & M. Lewis, 2013. Allee effect from parasite spill-back. American Naturalist. 182, 640-652.

35. *Thitiwan P., J. Sanchez, E.E. **Rees, M. Krkosek, S.R.M. Jones & C.W. Revie, 2013. Sea lice infestations on juvenile chum and pink salmon in the Broughton Archipelago, Canada from 2003 to 2012. Diseases of Aquatic Organisms. 105, 149-161.

34. *Rogers, L., S. *Peacock, P. McKenzie, S. DeDominicis, S. Jones, P. Chandler, M. Foreman, C. Revie, and M. Krkosek. Modeling parasite dynamics on farmed salmon for precautionary conservation management of wild salmon. PLoS ONE. 8: e60096.

33. *Senior, A., M. Krkosek, & S. Nakagawa. The practicality of Trojan sex chromosomes as a biological control: An agent based model of two highly invasive Gambusia species. Biological Invasions. In press.

32. *Peacock, S., M. Krkosek, S. Proboszcz, C. Orr, & M. Lewis, 2013. Cessation of a salmon decline with control of parasites. Ecological Applications. 23, 606-620.
News coverage by the Globe & Mail and CBC.

31. *Paul, S., M. Krkosek, K. Probert, & G. Closs, 2013. Osmoregulation and survival of two species of Tenagomysis (Crustacea: Mysidacea) in estuaries of southern New Zealand. Marine and Freshwater Research. 64, 340-347.

30. Krkosek, M., C. Revie, P. Gargan, O. Skilbrei, B. Finstad, & C. Todd, 2013. Impact of parasites on salmon recruitment in the Northeast Atlantic Ocean. Proceedings of the Royal Society B. 280, 20122359.
F1000 recommended and news coverage by BBC


29. **Connors, B., D. *Braun, R. Peterman, A. Cooper, J. Reynolds, L. Dill, G. Ruggerone, and M. Krkosek, 2012. Migration links ocean-scale competition and local ocean conditions with exposure to farmed salmon to shape wild salmon dynamics. Conservation Letters. 5, 304-312.
Cover story for the issue and news coverage in the Vancouver Sun.

28. Frazer, L.N., A. Morton, and M. Krkosek 2012. Host density thresholds in sea lice epidemics: Evidence, sensitivity, and subcritical estimation. Proceedings of the Royal Society B, 279, 1950-1958.

27. Krkosek, M., B. **Connors, M. Lewis, and R. Poulin. 2012. Allee effects may slow the spread of parasites in a coastal marine ecosystem. American Naturalist, 179, 401-412.

26. *Ashander, J., M. Krkosek, & M. Lewis. 2012. Aquaculture-induced changes to dynamics of a migratory host and specialist parasite: a case study of pink salmon and sea lice. Theoretical Ecology, 5, 231-252.


25. Krkosek, M., B. *Connors, A. Morton, M. Lewis, L. Dill, and R. Hilborn. 2011. Effects of parasites from salmon farms on productivity of wild salmon. Proceedings of the National Academy of Sciences of the USA. 108, 14700-14704.
News coverage by CBC.

24. Williams, R., M. **Krkosek, E. Ashe, T. A. **Branch, S. Clark, P. Hammond, E. *Hoyt, D. Noren, D. Rosen and A. Winship. 2011. Endangered killer whales and endangered salmon:  Conflicting objectives between conservation and fisheries. PLoS ONE, 6, e26738.
News coverage by Discovery Channel.

23. **Krkosek, M., R. Hilborn, R. Peterman, & T. Quinn. 2011. Cycles, stochastcity, and density dependence in pink salmon population dynamics. Proceedings of the Royal Society of London Series B, 278, 2060-2068.

23. **Krkosek, M., B. *Connors, P. *Mages, S. ^Peacock, H. ^Ford, J. *Ford, A. Morton, J. Volpe, R. Hilborn, L. Dill, M. Lewis. 2011. Fish farms, parasites, and predators: Implications for salmon population dynamics. Ecological Applications. 21, 897-914.

22. **Krkosek, M., & R. Hilborn. 2011. Sea lice infestations and productivity of pink salmon populations in the Broughton Archipelago. Canadian Journal of Fisheries and Aquatic Sciences. 68, 17-29.


21. *Krkosek, M., A. *Bateman, S. Proboscsz, & C. Orr. 2010. Dynamics of outbreak and control of salmon lice on two salmon farms is the Broughton Archipelago. Aquaculture Environment Interactions. 1, 137-146.

20. *Connors, B., M. *Krkosek, J. *Ford, & L. Dill. 2010. Coho salmon productivity in relation to direct and trophic transmission of sea lice from salmon aquaculture. Journal of Applied Ecology. 47, 1372-1377.

19. Lewis, M.A., M. *Krkosek, M.J.Wonham. 2010. Dynamics of emerging wildlife disease. In S. Sivaloganathan (Ed) Mathematical Biology, Fields Institute Communications, American Mathematical Society.

18. Morton, A., A. ^McConnell, R. Routledge, M. *Krkosek. 2010. Sea lice dispersion and salmon survival in relation to fallowing and chemical treatment on salmon farms. ICES Journal of Marine Science. 68, 144-156.

17. **Krkosek, M. 2010. Host density thresholds and disease control for fisheries and aquaculture. Aquaculture Environment Interactions. 1, 21-32.
Invited Article for 1st Issue

16. *Krkosek, M. 2010. Sea lice and salmon in Pacific Canada: Ecology and policy. Frontiers in Ecology and the Environment. 8, 201-209.

16. *Krkosek, M. and M. Lewis. 2010. An R0 theory for source-sink dynamics with application to Dreissena competition. Theoretical Ecology. 3, 24-43.


15. *Krkosek, M., A. Morton, J. Volpe, & M. Lewis. 2009. Sea lice and salmon population dynamics: Effects of exposure time for migratory fish. Proceedings of the Royal Society of London Series B. 276, 2819-2828.

14. Keller, R., M.A. Lewis, D.M. Lodge, J. F. Shogren & M. *Krkosek, 2009. Putting bioeconomic research into practice.  In Bioeconomics of Invasive Species: Integrating ecology, economics, policy and management. Editors: R. Keller, D.M. Lodge, M. A. Lewis and J. F. Shogren. Oxford University Press. Oxford.


13. *Krkosek, M., J. *Ford, A. Morton, S. Lele, M. Lewis, 2008. Response to comment on “Declining wild salmon populations in relation to parasites from farm salmon”. Science. 322, 1790-1791.

12. *Connors, B., M. *Krkosek, & L. Dill, 2008. Sea lice escape predation on their host. Biology Letters. 4, 455-457.
News coverage in the Vancouver Sun and by Science.

11. *Krkosek, M., J. *Ford, A. Morton, S. Lele, M. Lewis, 2008. Sea lice and pink salmon declines: response to Brooks and Jones. Reviews in Fisheries Science. 16, 413-420.
News coverage in the Vancouver Sun.

10. Morton, A., R. Routledge, & M. *Krkosek. 2008. Sea lice infestation of juvenile salmon and herring associated with fish farms off the east central coast of British Columbia. North American Journal of Fisheries Management. 28, 523-532.


9. *Krkosek, M., J. *Ford, A. Morton, S. Lele, R.A. Myers, M. Lewis, 2007. Declining wild salmon populations in relation to parasites from farm salmon. Science. 318, 1772-1775.
News coverage by Science, Nature, Economist, NY Times, CBC, BBC, Globe & Mail, others.

8. *Krkosek, M., A. Gottesfeld, B. Proctor, D. Rolston, C. Carr-Harris, M. Lewis, 2007. Effects of host migration, diversity, and aquaculture on sea lice threats to wild Pacific salmon populations. Proceedings of the Royal Society of London Series B. 274, 1341-3149.

7. *Krkosek, M., J.S. Lauzon-Guay, & M. Lewis. 2007. Relating dispersal and range expansion of California sea otters. Theoretical Population Biology. 71, 401-407.


6. *Krkosek, M., M. Lewis, A. Morton, L.N. Frazer & J. Volpe. 2006. Epizootics of wild fish induced by farm fish. Proceedings of the National Academy of Sciences of the USA. 103, 15506-15510.
F1000 recommended, editorial coverage in PNAS and news coverage.

5. *Krkosek, M., M. Lewis, J. Volpe, & A. Morton. 2006. Fish Farms and sea lice infestations of wild juvenile salmon in the Broughton Archipelago – A rebuttal to Brooks (2005). Reviews in Fisheries Science. 14: 1-11.


4. *Krkosek, M., M.A. Lewis, & J.P. Volpe. 2005. Transmission dynamics of parasitic sea lice from farm to wild salmon. Proceedings of the Royal Society of London Series B. 272, 689-696.
Cover story for the Proc B issue and news in NY Times, Globe & Mail, MacLeans, CBC (As It Happens), others.

3. *Krkosek, M., A. Morton, & J.P. Volpe. 2005. Non-lethal assessment of juvenile Pacific salmon for parasitic sea lice infections and fish health. Transactions of the American Fisheries Society. 134, 711-716.


2. Yang, K., D. Langlois, L. Campbell, J. Challis, M. Krkosek, & M.Yu, 1997. Cellular localization and developmental regulation of 11 beta-hydroxysteroid dehydrogenase type 1 (11 beta-HSD1) gene expression in the ovine placenta. Placenta. 18 503-509.

1. Yang, K., M. Fraser, M. Yu, M. Krkosek, J.R.G. Challis, G.E. Lamming, L.E. Campbell, & A. Darnel, 1996. Pattern of 11-beta-hydroxysteroid dehydrogenase type 1 messenger ribonucleic acid expression in the ovine uterus during the estrous cycle and pregnancy. Biology of Reproduction. 55 1231-1236.